| Preface | |
| Ch. 1 | Shells and the Questions of Biology | 3 |
| Pt. I | The Rules of Shell Construction | |
| Ch. 2 | Themes and Variations: The Geometry of Shells | 9 |
| Ch. 3 | The Economics of Construction and Maintenance | 39 |
| Pt. II | Life in a Dangerous World: How Shells Work | |
| Ch. 4 | The Mechanics of Shells | 61 |
| Ch. 5 | Predators and Their Methods | 94 |
| Ch. 6 | Coping with Enemies: The Shell as Protection | 113 |
| Pt. III | The Dimension of Time | |
| Ch. 7 | A Historical Geography of Shells | 153 |
| Ch. 8 | Evolutionary Economics: The Rise and Fall of Adaptive Themes | 174 |
| Index | 203 |